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The evil organization Shocker alters history by sending their newest creation, Kamen Rider 3, to defeat Kamen Riders 1 and 2 back in 1973. In the alternate present, Shocker is more powerful than ever, and most of the Kamen Riders are now either imprisoned or brainwashed to work for them. One such brainwashed Rider is Shinnosuke Tomari who thanks to his partner Kiriko Shijima, remembers the truth and joins forces with the remaining Kamen Riders who are still at large to make a stand against Shocker and put history back on its proper course.
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The Gokaigers were captured by the Zangyack remnants after departing to the Zangyack homeworld to destroy the empire once and for all. The Gokaigers, now working for the newly reformed Zangyack empire, return to Earth with a black Gokai Galleon and attack the Go-Busters in search of five phantom Ranger Keys which will unlock the "Greatest Power in the Universe". During the fight with the Gokaigers, the Go-Busters are able capture Navi, who tells them that the Zangyack are also working with Vaglass and that they must go back in time to save the other members of the Super Sentai Teams and defeat the alliance of Zangyack and Vaglass.
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The Underground Empire Badan makes its move, intending to invert the world. Kouta and Mai are pulled through one of these inversions where the discover a boy named Shu, who also holds the power to invert the world. As the Riders fight to protect Shu, Fifteen appears again, seeking Shu. As Kouta flees, past Riders gather and fight one another, split between the Showa and Heisei era Riders. But, the truth of Badan is not as it seems…