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The “Amazon Livestock Project,” a mysterious care facility, the Amazon extermination organization 4C (Competitive Creatures Control Center) pursuing Haruka Misuzawa . . . all come together in a tale of symbiosis and competition; the mortal combat between the carnivore and herbivore. And, finally, an end to the troubles and conflicts of the Riders, two beings confronting a fierce fate.
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Ashitaka, a prince of the disappearing Emishi people, is cursed by a demonized boar god and must journey to the west to find a cure. Along the way, he encounters San, a young human woman fighting to protect the forest, and Lady Eboshi, who is trying to destroy it. Ashitaka must find a way to bring balance to this conflict.
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Sakiko, a young bank teller, has an unhealthy obsession with money. Thieves hold up the bank, kidnapping Sakiko in the process, but eventually crash their car, resulting in a suitcase stuffed with cash falling into a nearby river. For the remainder of the film, Sakiko begins a desperate quest to retrieve the money.