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When an artifact known as the "King's Seal" is stolen during transport from Soul Society, Hitsugaya Toushirou is assigned to retrieve it. Toushirou goes missing after a battle with the thieves, leading Seireitei to suspect him of treachery. They order his immediate capture and execution. Unwilling to believe him capable of such a crime, Ichigo, Rangiku, Rukia, and Renji set out to find Toushirou.
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It is not strange that the Demon Lord's forces fear the Crimson Demons, the clan from which Megumin and Yunyun originate. Even if the Demon Lord's generals attack their village, the Crimson Demons can just easily brush them off with their supreme mastery of advanced and overpowered magic. When Yunyun receives a seemingly serious letter regarding a potential disaster coming to her hometown, she immediately informs Kazuma Satou and the rest of his party. After a series of wacky misunderstandings, it turns out to be a mere prank by her fellow demon who wants to be an author. Even so, Megumin becomes worried about her family and sets out toward the Crimson Demons' village with the gang. There, Kazuma and the others decide to sightsee the wonders of Megumin's birthplace. However, they soon come to realize that the nonsense threat they received might have been more than just a joke.
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The special takes place during the two year before the Straw Hats reunite on Sabody. Luffy is currently in Rusukaina training to get stronger to take on the New World. However the training is interrupted when Hancock's sisters, Marigold and Sandersonia, are kidnapped by the Byrnndi World, a pirate who was locked away on Level 6 of Impel Down but escaped during Luffy's invasion to save Ace, in order to lure Hancock to him and use her as a hostage against the World Government due to her Shichibukai status. Thus Luffy and Hancock head off to confront him and save Hancock's sisters.