The long form of the trope title is "Out of Character Is Serious Business." Some characters have strong traits that they are known by. This is for when characters momentarily break away from their normal habits to make a point about the seriousness of the situation. Often causes the other characters to do a Double Take and mention why this event is Serious Business. When most or all of these OoC moments happen at once, you can be sure that the Darkest Hour has arrived, leading characters to behave in ways they normally wouldn't, because they know they might not have another chance to do so.
Fan Casting
Ebenezer Scrooge (Scrooge 1970)
Bigwig (Watership Down)
Sergeant Arthur Wilson
Sonic the Hedgehog (Movie)
Dr. Coomer
Wakko Warner
Lunar Guy
Josh Nichols
Mandy Mason
Timmy Collins
Sam Barker
Benedick (Much Ado About Nothing)
Phineas Flynn
Isabella Garcia-Shapiro
MAX (Flight Of The Navigator)
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