In 1942, Captain America and Bucky Barnes were summoned by president Franklin D. Roosevelt to investigate a series of ships sinking in the Sargasso Sea, where they meet the Sub-Mariner, an Atlantean prince who vowed to wreak vengeance on all surface dwellers. Meanwhile, a scientist named Phineas T. Horton showed off to the press his latest invention: an android named the Human Torch. The fiery android escaped and adopted a young orphan named Thomas Raymond, who worked in a circus under the name of Toro.
Thru various circumstances, Captain America and Bucky alongside Namor the Sub-Mariner meets the Human Torch and Toro and team dup to fight the Red Skull and Arnim Zola and prevent them from invading Manhattan.